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Skeleton
Gist
A skeleton is the rigid framework of bones and cartilage that supports an organism's body, protects internal organs, and allows for movement; it can be internal (endoskeleton, like in humans and vertebrates) or external (exoskeleton, like in insects). The human skeleton, specifically, is an internal structure with 206 bones in adulthood, divided into the axial (skull, spine, ribs) and appendicular (limbs, girdles) parts, working with ligaments, tendons, and cartilage to provide shape, support, and facilitate motion.
A skeletal system is necessary to support the body, protect internal organs, and allow for the movement of an organism. There are three different skeleton designs that fulfill these functions: hydrostatic skeleton, exoskeleton, and endoskeleton.
Summary
A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal frame to which the organs and soft tissues attach; and the hydroskeleton, a flexible internal structure supported by the hydrostatic pressure of body fluids.
Vertebrates are animals with an endoskeleton centered around an axial vertebral column, and their skeletons are typically composed of bones and cartilages. Invertebrates are other animals that lack a vertebral column, and their skeletons vary, including hard-shelled exoskeleton (arthropods and most molluscs), plated internal shells (e.g. cuttlebones in some cephalopods) or rods (e.g. ossicles in echinoderms), hydrostatically supported body cavities (most), and spicules (sponges). Cartilage is a rigid connective tissue that is found in the skeletal systems of vertebrates and invertebrates.
Details
Human skeleton is the internal skeleton that serves as a framework for the body. This framework consists of many individual bones and cartilages. There also are bands of fibrous connective tissue—the ligaments and the tendons—in intimate relationship with the parts of the skeleton. This article is concerned primarily with the gross structure and the function of the skeleton of the normal human adult.
The human skeleton, like that of other vertebrates, consists of two principal subdivisions, each with origins distinct from the others and each presenting certain individual features. These are (1) the axial, comprising the vertebral column—the spine—and much of the skull, and (2) the appendicular, to which the pelvic (hip) and pectoral (shoulder) girdles and the bones and cartilages of the limbs belong. A third subdivision, the visceral (splanchnocranium), comprises the lower jaw, some elements of the upper jaw, and the branchial arches, including the hyoid bone.
When one considers the relation of these subdivisions of the skeleton to the soft parts of the human body—such as the nervous system, the digestive system, the respiratory system, the cardiovascular system, and the voluntary muscles of the muscle system—it is clear that the functions of the skeleton are of three different types: support, protection, and motion. Of these functions, support is the most primitive and the oldest; likewise, the axial part of the skeleton was the first to evolve. The vertebral column, corresponding to the notochord in lower organisms, is the main support of the trunk.
The central nervous system lies largely within the axial skeleton, the brain being well protected by the cranium and the spinal cord by the vertebral column, by means of the bony neural arches (the arches of bone that encircle the spinal cord) and the intervening ligaments.
A distinctive characteristic of humans as compared with other mammals is erect posture. The human body is to some extent like a walking tower that moves on pillars, represented by the legs. Tremendous advantages have been gained from this erect posture, the chief among which has been the freeing of the arms for a great variety of uses. Nevertheless, erect posture has created a number of mechanical problems—in particular, weight bearing. These problems have had to be met by adaptations of the skeletal system.
Protection of the heart, lungs, and other organs and structures in the chest creates a problem somewhat different from that of the central nervous system. These organs, the function of which involves motion, expansion, and contraction, must have a flexible and elastic protective covering. Such a covering is provided by the bony thoracic basket, or rib cage, which forms the skeleton of the wall of the chest, or thorax. The connection of the ribs to the breastbone—the sternum—is in all cases a secondary one, brought about by the relatively pliable rib (costal) cartilages. The small joints between the ribs and the vertebrae permit a gliding motion of the ribs on the vertebrae during breathing and other activities. The motion is limited by the ligamentous attachments between ribs and vertebrae.
The third general function of the skeleton is that of motion. The great majority of the skeletal muscles are firmly anchored to the skeleton, usually to at least two bones and in some cases to many bones. Thus, the motions of the body and its parts, all the way from the lunge of the football player to the delicate manipulations of a handicraft artist or of the use of complicated instruments by a scientist, are made possible by separate and individual engineering arrangements between muscle and bone.
Additional Information
The skeleton is a remarkable organ that provides the body with a frame that is strong enough for protection, light enough for mobility, and adaptable for changing structural needs. The skeleton also serves metabolic functions as a storehouse for calcium and phosphorus, a buffering site for hydrogen ion excess, and a binding site for toxic ions such as lead and aluminum. When skeletal tissues are required to fulfill these latter functions, this may occur at the cost of structural integrity and lead to fractures. Once the adult skeleton has been formed, both the structural and metabolic functions are carried out largely by remodeling—removal and replacement of bone tissue at the same site in so-called bone multicellular units (BMU)—rather than modeling, which is formation of bone at sites where no prior resorption has occurred. Both processes do continue throughout life, however. In particular, modeling in the form of new periosteal apposition can occur with aging as a compensatory mechanism to the weakening of bone by the trabecular and endosteal loss and cortical porosity that occurs with increased resorption and inadequate formation in BMUs.

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