Math Is Fun Forum

Jai Ganesh

Administrator

Registered: 2005-06-28

Posts: 48,384

Apiculture

Apiculture

Gist

Apiculture is the science of raising or maintaining colonies of bees and their hives (beekeeping).

Apiculture - the science and art of raising honey bees. Apis mellifera - scientific name of the honey bee found in the United States. Automatic uncapper - automated device that removes the cappings from honey combs, usually by moving heated knives, metal teeth, or flails.

Summary

Archaeological evidence shows us that the relationship between humans and bees is at least 17,000 years old, with the earliest depictions of human honey hunting and harvesting of bee products occurring 4,000 years before our earliest systems of domesticated agriculture; but it wasn’t until nearly 14,000 years later that we find the first evidence of sophisticated bee colony management as part of an agricultural system.

With more than 4,000 native bees in North America, honey bees take much of the spotlight in agriculture - but they are not native to our country. They were imported from Europe in the 17th century.

Details

Beekeeping (or apiculture) is the maintenance of bee colonies, commonly in artificial beehives. Honey bees in the genus Apis are the most commonly kept species but other honey producing bees such as Melipona stingless bees are also kept. Beekeepers (or apiarists) keep bees to collect honey and other products of the hive: beeswax, propolis, bee pollen, and royal jelly. Other sources of beekeeping income include pollination of crops, raising queens, and production of package bees for sale. Bee hives are kept in an apiary or "bee yard".

The keeping of bees by humans, primarily for honey production, began around 10,000 years ago. A sample of 5,500-year-old honey was unearthed from the grave of a noblewoman during archaeological excavations in 2003 near the town of Borjomi, Georgia. Ceramic jars found in the grave contained several types of honey, including linden and flower honey. Domestication of bees can be seen in Egyptian art from around 4,500 years ago; there is also evidence of beekeeping in ancient China, Greece, and Maya.

In the modern era, beekeeping is often used for crop pollination and the collection of byproducts, such as wax and propolis. The largest beekeeping operations are agricultural businesses but many small beekeeping operations are run as a hobby. As beekeeping technology has advanced, beekeeping has become more accessible, and urban beekeeping was described as a growing trend as of 2016. Some studies have found city-kept bees are healthier than those in rural settings because there are fewer pesticides and greater biodiversity in cities.

History:

Early history

At least 10,000 years ago, humans began to attempt to maintain colonies of wild bees in artificial hives made from hollow logs, wooden boxes, pottery vessels, and woven straw baskets known as skeps. Depictions of humans collecting honey from wild bees date to 10,000 years ago. Beekeeping in pottery vessels began about 9,000 years ago in North Africa. Traces of beeswax have been found in potsherds throughout the Middle East beginning about 7,000 BCE. Domestication of bees is shown in Egyptian art from around 4,500 years ago. Simple hives and smoke were used, and honey was stored in jars, some of which were found in the tombs of pharaohs such as Tutankhamun. In the 18th century, European understanding of the colonies and biology of bees allowed the construction of the movable comb hive so honey could be harvested without destroying the entire colony.

Honeybees were kept in Egypt from antiquity. On the walls of the sun temple of Nyuserre Ini from the Fifth Dynasty before 2,422 BCE, workers are depicted blowing smoke into hives as they remove honeycombs. Inscriptions detailing the production of honey are found on the tomb of Pabasa from the Twenty-sixth Dynasty c. 650 BCE, in which cylindrical hives are depicted along with people pouring honey into jars.

An inscription records the introduction of honey bees into the land of Suhum in Mesopotamia, where they were previously unknown:

I am Shamash-resh-ușur, the governor of Suhu and the land of Mari. Bees that collect honey, which none of my ancestors had ever seen or brought into the land of Suhu, I brought down from the mountain of the men of Habha, and made them settle in the orchards of the town 'Gabbari-built-it'. They collect honey and wax, and I know how to melt the honey and wax – and the gardeners know too. Whoever comes in the future, may he ask the old men of the town, (who will say) thus: "They are the buildings of Shamash-resh-ușur, the governor of Suhu, who introduced honey bees into the land of Suhu".

— translated text from Stele, (Dalley, 2002)

The oldest archaeological finds directly relating to beekeeping have been discovered at Rehov, a Bronze and Iron Age archaeological site in the Jordan Valley, Israel. Thirty intact hives made of straw and unbaked clay were discovered in the ruins of the city, dating from about 900 BCE, by archaeologist Amihai Mazar. The hives were found in orderly rows, three high, in a manner that according to Mazar could have accommodated around 100 hives, held more than one million bees and had a potential annual yield of 500 kilograms (1,100 lb) of honey and 70 kilograms (150 lb) of beeswax, and are evidence an advanced honey industry in Tel Rehov, Israel 3,000 years ago.

In ancient Greece, in Crete and Mycenae, there existed a system of high-status apiculture that is evidenced by the finds of hives, smoking pots, honey extractors and other beekeeping paraphernalia in Knossos. Beekeeping was considered a highly valued industry controlled by beekeeping overseers—owners of gold rings depicting apiculture scenes rather than religious ones as they have been reinterpreted recently, contra Sir Arthur Evans. Aspects of the lives of bees and beekeeping are discussed at length by Aristotle. Beekeeping was also documented by the Roman writers Virgil, Gaius Julius Hyginus, Varro, and Columella.

Beekeeping has been practiced in ancient China since antiquity. In a book written by Fan Li (or Tao Zhu Gong) during the Spring and Autumn period are sections describing beekeeping, stressing the importance of the quality of the wooden box used and its effects on the quality of the honey.[19] The Chinese word for honey (mì, reconstructed Old Chinese pronunciation *mjit) was borrowed from proto-Tocharian *ḿət(ə) (where *ḿ is palatalized; cf. Tocharian B mit), cognate with English mead.

The ancient Maya domesticated a species of stingless bee, which they used for several purposes, including making balché, a mead-like alcoholic drink. By 300 BCE they had achieved the highest levels of stingless beekeeping practices in the world. The use of stingless bees is referred to as meliponiculture, which is named after bees of the tribe Meliponini such as Melipona quadrifasciata in Brazil. This variation of beekeeping still occurs today. For instance, in Australia, the stingless bee Tetragonula carbonaria is kept for the production of honey.

Scientific study of honey bees

European natural philosophers began to scientifically study bee colonies in the 18th century. Eminent among these scientists were Swammerdam, René Antoine Ferchault de Réaumur, Charles Bonnet and François Huber. Swammerdam and Réaumur were among the first to use a microscope and dissection to understand the internal biology of honey bees. Réaumur was among the first to construct a glass-walled observation hive to better observe activities inside hives. He observed queens laying eggs in open cells but did not know how queens were fertilized; the mating of a queen and drone had not yet been observed and many theories held queens were "self-fertile" while others believed a vapor or "miasma" emanating from the drones fertilized queens without physical contact. Huber was the first to prove by observation and experiment that drones physically inseminate queens outside the confines of the hive, usually a great distance away.

Following Réaumur's design, Huber built improved glass-walled observation hives and sectional hives that could be opened like the leaves of a book. This allowed the inspection of individual wax combs and greatly improved direct observation of hive activity. Although he went blind before he was twenty, Huber employed a secretary named François Burnens to make daily observations, conduct experiments, and keep accurate notes for more than twenty years. Huber confirmed a hive consists of one queen, who is the mother of every female worker and male drone in the colony. He was also the first to confirm mating with drones takes place outside hives and that queens are inseminated in successive matings with male drones, which occur high in the air at a great distance from the hive. Together, Huber and Burnens dissected bees under the microscope, and were among the first to describe the ovaries and spermatheca (sperm store) of queens, as well as the math of male drones. Huber is regarded as "the father of modern bee-science" and his work Nouvelles Observations sur Les Abeilles (New Observations on Bees) revealed all of the basic scientific facts of the biology and ecology of honeybees.

Hive designs

Before the invention of the movable comb hive, the harvesting of honey frequently resulted in the destruction of the whole colony. The wild hive was broken into using smoke to quieten the bees. The honeycombs were pulled out and either immediately eaten whole or crushed, along with the eggs, larvae, and honey they held. A sieve or basket was used to separate the liquid honey from the demolished brood nest. In medieval times in northern Europe, although skeps and other containers were made to house bees, the honey and wax were still extracted after the bee colony was killed. This was usually accomplished by using burning sulfur to suffocate the colony without harming the honey within. It was impossible to replace old, dark-brown brood comb in which larval bees are constricted by layers of shed pupal skins.

The movable frames of modern hives are considered to have been developed from the traditional basket top bar (movable comb) hives of Greece, which allowed the beekeeper to avoid killing the bees. The oldest evidence of their use dates to 1669, although it is probable their use is more than 3,000 years old.

Intermediate stages in the transition from older methods of beekeeping were recorded in 1768 by Thomas Wildman, who described advances over the destructive, skep-based method so bees no longer had to be killed to harvest their honey. Wildman fixed an array of parallel wooden bars across the top of a straw hive 10 inches (25 cm) in diameter "so that there are in all seven bars of deal to which the bees fix their combs", foreshadowing future uses of movable-comb hives. He also described using such hives in a multi-story configuration, foreshadowing the modern use of supers: he added successive straw hives below and later removed the ones above when free of brood and filled with honey so the bees could be separately preserved at the harvest the following season. Wildman also described the use of hives with "sliding frames" in which the bees would build their comb.

Wildman's book acknowledges the advances in knowledge of bees made by Swammerdam, Maraldi, and de Réaumur—he includes a lengthy translation of Réaumur's account of the natural history of bees. Wildman also describes the initiatives of others in designing hives for the preservation of bees when taking the harvest, citing reports from Brittany in the 1750s due to the Comte de la Bourdonnaye. Another hive design was invented by Rev. John Thorley in 1744; the hive was placed in a bell jar that was screwed onto a wicker basket. The bees were free to move from the basket to the jar, and honey was produced and stored in the jar. The hive was designed to keep the bees from swarming as much as they would have in other hive designs.

In the 19th century, changes in beekeeping practice were completed through the development of the movable comb hive by the American Lorenzo Lorraine Langstroth, who was the first person to make practical use of Huber's earlier discovery of a specific spatial distance between the wax combs, later called the bee space, which bees do not block with wax but keep as a free passage. Having determined this bee space, which is commonly given as between 6 and 9 mm (0.24 and 0.35 in), though up to 15 mm (0.59 in) has been found in populations in Ethiopia. Langstroth then designed a series of wooden frames within a rectangular hive box, carefully maintaining the correct space between successive frames. He found the bees would build parallel honeycombs in the box without bonding them to each other or to the hive walls. This enables the beekeeper to slide any frame out of the hive for inspection without harming the bees or the comb; and protecting the eggs, larvae and pupae in the cells. It also meant combs containing honey could be gently removed and the honey extracted without destroying the comb. The emptied honeycombs could then be returned intact to the bees for refilling. Langstroth's book The Hive and Honey-bee (1853), describes his rediscovery of the bee space and the development of his patent movable comb hive. The invention and development of the movable comb hive enabled the growth of large-scale, commercial honey production in both Europe and the U.S.

20th and 21st century hive designs

Langstroth's design of movable comb hives was adopted by apiarists and inventors in both North America and Europe, and a wide range of moveable comb hives were developed in England, France, Germany and the United States. Classic designs evolved in each country; Dadant hives and Langstroth hives are still dominant in the U.S.; in France the De-Layens trough hive became popular, in the UK a British National hive became standard by the 1930s, although in Scotland the smaller Smith hive is still popular. In some Scandinavian countries and in Russia, the traditional trough hive persisted until late in the 20th century and is still kept in some areas. The Langstroth and Dadant designs, however, remain ubiquitous in the U.S. and in many parts of Europe, though Sweden, Denmark, Germany, France and Italy all have their own national hive designs. Regional variations of hive were developed according to climate, floral productivity and reproductive characteristics of the subspecies of native honey bees in each bio-region.

The differences in hive dimensions are insignificant in comparison to the common factors in these hives: they are all square or rectangular; they all use movable wooden frames; and they all consist of a floor, brood-box, honey super, crown-board and roof. Hives have traditionally been constructed from cedar, pine or cypress wood but in recent years, hives made from injection-molded, dense polystyrene have become increasingly common. Hives also use queen excluders between the brood-box and honey supers to keep the queen from laying eggs in cells next to those containing honey intended for consumption. With the 20th-century advent of mite pests, hive floors are often replaced, either temporarily or permanently, with a wire mesh and a removable tray.

In 2015, the Flow Hive system was invented in Australia by Cedar Anderson and his father Stuart Anderson, whose design allows honey to be extracted without cumbersome centrifuge equipment.

Pioneers of practical and commercial beekeeping

In the 19th century, improvements were made in the design and production of beehives, systems of management and husbandry, stock improvement by selective breeding, honey extraction and marketing. Notable innovators of modern beekeeping include:

Prokopovych's beehive system

Petro Prokopovych used frames with channels in the side of the woodwork; these were packed side-by-side in stacked boxes. Bees traveled between frames and boxes via these channels, which were similar to the cutouts in the sides of modern wooden sections.

Jan Dzierżon' beehive design has influenced modern beehives.

François Huber made significant discoveries about the bee life cycle and communication between bees. Despite being blind, Huber discovered a large amount of information about the queen bee's mating habits and her contact with the rest of the hive. His work was published as New Observations on the Natural History of Bees.

L. L. Langstroth has influenced modern beekeeping practice more than anyone else. His book The Hive and Honey-bee was published in 1853.

Moses Quinby, author of Mysteries of Bee-Keeping Explained, invented the bee smoker in 1873.

Amos Root, author of the A B C of Bee Culture, which has been continuously revised and remains in print, pioneered the manufacture of hives and the distribution of bee packages in the United States.

A. J. Cook author of The Bee-Keepers' Guide; or Manual of the Apiary, 1876.

Dr. C.C. Miller was one of the first entrepreneurs to make a living from apiculture. By 1878, he made beekeeping his sole business activity. His book, Fifty Years Among the Bees, remains a classic and his influence on bee management persists into the 21st century.

Franz Hruschka was an Austrian/Italian military officer who in 1865 invented a simple machine for extracting honey from the comb by means of centrifugal force. His original idea was to support combs in a metal framework and then spin them within a container to collect honey that was thrown out by centrifugal force. This meant honeycombs could be returned to a hive empty and undamaged, saving the bees a vast amount of work, time and materials. This invention significantly improved the efficiency of honey harvesting and catalyzed the modern honey industry.

Walter T. Kelley was an American pioneer of modern beekeeping in the early-and mid-20th century. He greatly improved upon beekeeping equipment and clothing, and went on to manufacture these items and other equipment. His company sold products worldwide and his book How to Keep Bees & Sell Honey, encouraged a boom in beekeeping following World War II.

Cary W. Hartman (1859–1947), lecturer, well known beekeeping enthusiast and honey promoter was elected President of the California State Beekeepers' Association in 1921.

In the UK, practical beekeeping was led in the early 20th century by a few men, pre-eminently Brother Adam and his Buckfast bee, and R.O.B. Manley, author of books including Honey Production in the British Isles and inventor of the Manley frame, which is still universally popular in the UK. Other notable British pioneers include William Herrod-Hempsall and Gale.

Ahmed Zaky Abushady (1892–1955) was an Egyptian poet, medical doctor, bacteriologist, and bee scientist, who was active in England and Egypt in the early twentieth century. In 1919, Abushady patented a removable, standardized aluminum honeycomb. In the same year, he founded The Apis Club in Benson, Oxfordshire, which later became the International Bee Research Association (IBRA). In Egypt in the 1930s, Abushady established The Bee Kingdom League and its organ The Bee Kingdom.

Additional Information

Beekeeping is care and management of colonies of honeybees. They are kept for their honey and other products or their services as pollinators of fruit and vegetable blossoms or as a hobby. The practice is widespread: honeybees are kept in large cities and villages, on farms and rangelands, in forests and deserts, from the Arctic and Antarctic to the Equator. Honeybees are not domesticated. Those living in a man-made domicile called a beehive or hive are no different from those living in a colony in a tree.

In antiquity people knew that bees produce delicious honey, that they sting, and that they increase their numbers by swarming. By the 17th century they had learned the value of smoke in controlling them and had developed the screen veil as protection against stings. From the 17th to the 19th century, the key discoveries upon which modern beekeeping is founded were made. These included the mystery of the queen bee as the mother of nearly all the occupants of the hive, her curious mating technique, parthenogenetic development, the movable frame hives, and the fact that bees rear a new queen if the old one disappears.

Given this knowledge, people were able to divide a colony instead of relying on natural swarming. Then the development of the wax-comb foundation, the starter comb on which bees build straight, easily handled combs, and the discovery that honey can be centrifuged or extracted from them and the combs reused, paved the way for large-scale honey production and modern commercial beekeeping. The identification of bee diseases and their control with drugs, the value of pollen and pollen substitutes in producing strong colonies, and the artificial insemination of queens have increased the honey-production efficiency of colonies.

---

It appears to me that if one wants to make progress in mathematics, one should study the masters and not the pupils. - Niels Henrik Abel.

Nothing is better than reading and gaining more and more knowledge - Stephen William Hawking.